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SNAKE
- For other uses, see Snake (disambiguation).
- Ophidian redirects here. For the collectible card game, see Ophidian 2350.
Snakes (from Old English snaca, and ultimately from the Proto-Indo-European base *snag- or *sneg-, "to crawl"), also known as ophidians, are cold-blooded legless reptiles closely related to lizards, which share the order Squamata. There are also several species of legless lizard which superficially resemble snakes, but are not otherwise related to them. A love of snakes is called ophiophilia, a fear of snakes is called ophidiophobia. A specialist in snakes is an ophiologist.
An old synonym for snake is serpent (which comes from Old French, and ultimately from *serp-, "to creep"[1]); in modern usage this usually refers to a mythic or symbolic snake, and information about such creatures can be found under serpent (symbolism). This article deals with the biology of snakes.
[edit] Evolution
The phylogeny of snakes is poorly known due to the fact that snake skeletons are typically small and fragile, making fossilization unlikely. It has however been generally agreed, on the basis of morphology, that snakes descended from lizard-like ancestors. Recent research based on genetics and biochemistry confirms this; snakes form a venom clade with several extant lizard families.
Recent fossil evidence suggests that snakes directly evolved from burrowing lizards, either varanids or some other group. An early fossil snake, Najash rionegrina, was a two-legged burrowing animal with a sacrum, fully terrestrial. One extant analog of these putative ancestors is the earless monitor Lanthanotus of Borneo, although it also is semi-aquatic. As these ancestors became more subterranean, they lost their limbs and became more streamlined for burrowing. Features such as the transparent, fused eyelids and loss of external ears, according to this hypothesis, evolved to combat subterranean conditions (scratched corneas, dirt in the ears). According to this hypothesis, snakes re-emerged onto the surface of the land much as they are today. Other primitive snakes are known to have possessed hindlimbs but lacked a direct connection of the pelvic bones to the vertebrae, including Haasiophis, Pachyrhachis and Eupodophis) which are slightly older than Najash.
Modern boas do have vestigal hind limbs, tiny, clawed digits known as anal spurs and used to grasp during mating.
The alternative hypothesis, based on morphology, suggests that ancestors were related to mosasaurs — extinct aquatic reptiles from the Cretaceous — which in turn are thought to have derived from varanid lizards. Under this hypothesis, the fused, transparent eyelids of snakes are thought to have evolved to combat marine conditions (corneal water loss through osmosis), while the external ears were lost through disuse in an aquatic environment, ultimately leading to an animal similar in appearance to sea snakes of today. In the Late Cretaceous, snakes re-colonized the land much like they are today. Fossil snake remains are known from early Late Cretaceous marine sediments, which is consistent with this hypothesis, particularly as they are older than the terrestrial Najash rionegrina. Similar skull structure; reduced/absent limbs; and other anatomical features found in both mosasaurs and snakes lead to a positive cladistical correlation, though some features are also shared with varanids. Supposedly similar locomotion for both groups is also used as support for this hypothesis. Genetic studies have indicated that snakes are not especially related to monitor lizards, and (it has been claimed) therefore not to mosasaurs, the proposed ancestor in the aquatic scenario of their evolution. However, there is more evidence linking mosasaurs to snakes than to varanids. Fragmentary remains that have been found from the Jurassic and Early Cretaceous indicate deeper fossil records for these groups, which may eventually refute either hypothesis.
The great diversity of modern snakes appeared in the Paleocene, probably correlated with the adaptive radiation of mammals following the extinction of the dinosaurs.
All snakes are carnivorous, eating small animals including lizards and other snakes, rodents and other small mammals, birds, eggs or insects. Some snakes have a venomous bite, which they do use to kill their prey before eating it. Other snakes kill their prey by constriction. Still others swallow their prey whole and alive. Most snakes are very easy to feed in captivity, apart from a minority of species.
Snakes do not chew their food and have a very flexible lower jaw, the two halves of which are not rigidly attached, and numerous other joints in their skull (see snake skull), allowing them to open their mouths wide enough to swallow their prey whole, even if it is larger in diameter than the snake itself. It is a common misconception that snakes actually dislocate their lower jaw to consume large prey.
After eating, snakes become torpid while the process of digestion takes place. Digestion is an intensive activity, especially after the consumption of very large prey. In species which feed only sporadically, the entire intestine enters a reduced state between meals to conserve energy, and the digestive system is 'up-regulated' to full capacity within 48 hours of prey consumption. So much metabolic energy is involved in digestion that in Crotalus durissus, the Mexican rattlesnake, an increase of body temperature to as much as 14 degrees Celsius above the surrounding environment has been observed.[2]
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